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Meredith Johnson is a Ph.D. student in Arizona State University's School of Life Sciences. Meredith is studying the thermal biology of a native Sonoran Desert bee species that is the primary pollinator of three species of palo verde trees, crucial plant species in upland Sonoran Desert habitats. Her work will help us to understand the impact of climate change on the health of this important Sonoran Desert pollinator.


Brittany Burgard is a Master's degree student in Arizona State University's School of Life Sciences. Brittany is studying the effects of wildfire on plant communities in the Sonoran Desert. As native Sonoran Desert plants are not adapted to wildfire, Brittany will document the changes in these plant communities after wildfires, comparing the distribution of plants in burned and unburned areas.


Honorable Mention. Cassidy Kyler is a senior at Arizona State University majoring in Sustainability. Concerned with the decline of burrowing owl populations in Arizona, Cassidy's project involves creating a safe habitat for burrowing owls on the ASU Polytechnic campus. The project will contribute to the conservation education of the public, as visitors will be welcome to observe the owls on a public walking trail.


FRIENDS OF THE SONORAN DESERT (FSD) is pleased to invite applications for our student grant program. Grants will be awarded to students for proposals that will increase our understanding of the Sonoran Desert ecosystem and facilitate our mission to protect and conserve the Sonoran Desert throughout its range.

Qualified applicants include advanced undergraduate students and graduate students currently enrolled in a university. In 2022, two awards will be made of $500 each.

Criteria used to evaluate applications:

     1. Overall quality of the proposed project

     2. Match of the proposal with FSD ongoing programs

    The proposed project can be part of a larger student-driven project (honors thesis; MS thesis; etc.).               

    Applicants should make clear how FSD funds will advance the completion of their project.

Applications must include:

     1. Title of project

     2. 500 word summary of proposed project

     3. Project budget and brief justification

     4. Two letters of reference sent to the email address below. One letter must be from the student's academic supervisor. 

Grant Program Timeline:

     Call for proposals: 2 February 2022

     Deadline for proposal submission: 15 March 2022

     Award notification: 31 March 2022

     Short update of findings due: 15 September 2022

     Final report due: 1 December 2022

Short Update and Final Reports

     Grant recipients will submit a short update on their findings, a final project report, and 3 high-quality images     

     (JPEGs) with permission for FSD to use these photos to promote their work: one image will be of the student

     investigator, preferably in the field (required); and two from the following list, as applicable: the species being     

     studied; the​ habitat in which the study was conducted; workshop; product produced; etc.


Student Grant Application

PAGE 1 (of 2)

Applicant Name:  

Affiliation (university/department):  

Grade Level:  

Contact Information – e-mail address and phone number:  

Reference 1:  Supervisor's name:  

Affiliation (university/department):  

Contact Information – e-mail address and phone number:  

Reference 2:  Name:  

Affiliation (university/department):  

Contact Information – e-mail address and phone number: 

Project Title:  

Budget plus justification of expenses (use remainder of Page 1):

     Budget guidelines: no salaries allowed.  Allowed expenses include $15/day per diem;       

     $0.25/mile mileage allowance; supplies and minor equipment.

PAGE 2 (of 2)

Project Description:

     500 words maximum/no longer than 1 page

     Single spaced; include up to 3 relevant citation references.

     Address overarching issue, how you will address the issue, and relevance of the issue to the     

     goals of FSD.



Iris Garthwaite studies how environmental factors can limit the fitness and distribution of plants. Her study species is the Fremont cottonwood (Populus fremontii), an important but threatened tree native to southwestern riparian ecosystems. Recent studies show that climate change is likely to differentially impact genetically distinct populations of Fremont’s cottonwood.


Iris utilizes common garden experiments in which genetically distinct strains of a species are grown under identical environmental conditions. This protocol allows investigators to disentangle the effects of genetic and environmental variation on the growth pattern across the range of a species. Because plants must have water to survive, Iris focuses on the veins in leaves which are the transporters of water and nutrients. The architecture of leaf veins determines the ability of plants to transfer water, which in turn affects overall plant fitness.


Iris found that leaf-vein architecture varied significantly across a gradient of climates from which her common gardens were established – ranging from sites with mean annual temperatures of 10.7 – 22.8 degrees C. Furthermore, the magnitude of leaf-trait adaptability differed among populations of Fremont’s cottonwood.


The results of this study suggest that, to conserve this threatened riparian tree species,  managers engaged in ecological restoration could improve conservation project outcomes by choosing cuttings of Fremont’s cottonwood that are most resilient in response to climate change.

Sofia Salazar studies the species richness of oribatid mites, small organisms related to spiders, scorpions, and ticks, which are the most abundant and diverse group of small animals inhabiting soils. Oribatids play a key role in global biogeochemical cycles, biological regulation, and soil aeration.


Sophia collects soil oribatids in the Madrean Sky Islands of southern Arizona. While soil communities are recognized as a critical component of the biosphere, the below-ground biodiversity in this region, particularly the diversity of oribatids, is poorly known. To date, Sofia has collected 366 soil samples from two mountain ranges (Santa Rita and Dragoon) from which she has identified 8,557 adult oribatid mites belonging to 116 morphospecies (a morphospecies is defined by its most easily observable physical characters). Sofia has classified these morphospecies into 40 families and 25 superfamilies,17 of which are new records for Arizona. Sofia extracts DNA from each morphospecies, and these data are then uploaded as “barcodes” into an international database that will facilitate species identification and comparison of her data with other species.


Sofia is also examining the dispersal advantages of asexual and sexual oribatid species, determining the genetic structure of oribatid populations, and analyzing the relationship between species diversity and environmental and altitudinal variables. As many species of oribatids are used as indicators of overall soil biodiversity and health, becoming familiar with these organisms will help us more effectively conserve the fragile soils of the Sonoran Desert and its sky islands.



Sofia Salazar is a Ph.D. student in Arizona State University's School of Life Sciences. She was awarded a $500 student grant for her project to identify and describe the oribatid mites that inhabit the Madrean Sky Islands in southern Arizona. Sofia's project will help us to understand the vast underground biodiversity in Arizona's southern mountains, and will inform our advocacy for protecting soil habitats in the Sonoran Desert.

Iris Garthwaite is a Master's student in the Northern Arizona University's School of Earth and Sustainability. She was awarded a $500 student grant for her project that will use innovative methods to assess the adaptability of the Fremont cottonwood tree, an important and widespread species in riparian areas in the Sonoran Desert, to the impacts of climate change.



Miguel Grageda studies endangered Sonoran pronghorn in the Pinacate Biosphere Reserve in Sonora, Mexico, to see how nearby human development (roads, other barriers, and livestock grazing) might affect their population. 

Miguel's 81 camera traps cover 200,000 acres, a significant portion of the pronghorn habitat within the Reserve. He recorded an increase in camera-trap records (each time an individual or a group of individuals was obtained at a specific time and location) from 123 in 2019 to 137 in 2020. The sex ratio was approximately two males for every female. 

One of Miguel's most exciting findings for 2020 was a group of fawns born during the first week of March, demonstrating recruitment of pronghorn into this area. These animals were also recorded in September and October, thus they survived the harsh summer in the region. Overall, six independent records of fawns were recorded in 2019, and 11 in 2020.

While pronghorn were distributed throughout the Reserve, 37% of records (51/137) were made at four camera-trap locations along Highway 2, in proximity to the international border. The majority of these records (92%) were made at a distance of either 750 m or 1500 m from the highway, rather than close to this disturbance.​ These records were made in February, May, and November 2020, suggesting that pronghorn were present in the area for most of the year.

Miguel plans to collect data throughout 2021 in order to compare his findings over a three-year time span.


Julie Rakes studies biocrusts, the topsoil microbial communities found atop desert soils. It is important to understand what affects the health of biocrusts as they provide nutrients to soils in nutrient-poor regions such as the Sonoran Desert.


Recent field surveys of biocrusts have revealed the presence of virulent predatory bacteria (the “plaques” seen in Julie's photo), which infect healthy biocrusts. Julie's research involves analyzing the effects of rainfall and temperature on the health of biocrusts in desert soils.


Julie's data show that increased rainfall facilitates the rate of infection by predatory bacteria, killing up to 28% of the biocrusts that they inhabit. Intense sunlight has the opposite effect, killing about half of the infectious bacteria on the soil surface.


By learning more about what factors influence the health of biocrusts, we will be able to develop better management plans to protect them.



Julie Bethany Rakes is a Ph.D. student in ASU's School of Life Sciences. She was awarded a $500 grant to help her complete her studies on a predatory bacterium found in the the soil crusts of the Sonoran Desert. Julie's work will increase our understanding of how desert microbes impact topsoil communities in arid, desert environments.


Miguel Angel Grageda Garcia is a Ph.D. student in the University of Arizona's School of Natural Resources. He was awarded a $500 grant to continue his camera trap studies on the endangered Sonoran pronghorn in the El Pinacate Biosphere Reserve in Sonora, Mexico. Miguel is examining how obstacles like roads and physical barriers affect the movements and range of Sonoran pronghorn, as well as their ability to compete with livestock for food and space.